Don’t know if you solved this but it just happend to me after verifying links in Expressions 4. It added a _vti_conf folder to every folder including the Themes folder. Once I found the right folder and deleted it, the site functioned as normal.
Hope that helps!
This happened to me this week after opening the site live in MS Expressions 4 and asking it to verify backlinks. Didn’t save anything but it added the _vti_cnf folder to every folder. Deleted a number of them before I found the right one. It was in one of the folders relating to Themes. Once I deleted the correct one, the site started working like normal.
I learned my lesson. I was up until 2:00 am on a Sunday night trying to fix it!
Good luck.