Not exactly a plugin but a free theme which allows your users to post from the front-end. You could customize the theme to taste or check the code and apply to your own theme. http://wordpress.org/extend/themes/p2
Based on your initial message I would probably suggest taking a look at Buddypress though http://buddypress.org/. This relies on installing WordPress Multi User, and it will give the majority of the functionality you are after, with user profiles etc.
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